As described by Paine (1966), some sea stars (e.g., Pisaster ochraceus) may prey on sea urchins, mussels, and other shellfish that have no other natural predators. If the sea star is removed from the ecosystem, the mussel population explodes uncontrollably, driving out most other species. This means that sea stars are the _____ species in these ecosystems.单项选择题

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